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Spent two years as a Naval officer on the USS Manatee before returning to Stanford to attend graduate school in marine biology. BA and MA degrees in Biological Science from Stanford University, Ph.D. degree from the University of California at Berkeley.
Postdoctoral research in cell biology at the Laboratory for Quantitative Biology, University of Miami, Coral Gables, Florida. Taught invertebrate zoology, marine biology and marine ecology at the University of California at Santa Barbara, Oregon State University's Marine Science Center in Newport, Oregon, the Bodega Marine Laboratory of the University of California, and the Friday Harbor Laboratories of the University of Washington. Joined the Department of Biological Science at Florida State University as an assistant professor and retired as a Professor Emeritus of Biological Science. Presently residing in Tallahassee, Florida.